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Hub Rank	Node	Children	Parents	Degree	Betweenness
#1	ETS1	49	3	52	227.071
#2	FOXA1	47	3	50	93.814
#3	SPDEF	43	3	46	1860.62
#4	CREB3L4	39	3	42	1120.685
#5	HMGA2	38	8	46	708.056
#6	ELK3	36	1	37	55.696
#7	HOXA3	35	9	44	327.688
#8	XBP1	33	3	36	36.495
#9	AR	33	5	38	144.814
#10	DLX6	31	8	39	4508.812
#11	FOSL1	31	6	37	508.797
#12	ZEB1	30	6	36	511.214
#13	PBX1	30	8	38	3229.851
#14	ESR1	29	7	36	703.422
#15	PRDM1	28	9	37	1097.063
#16	GRHL2	28	4	32	139.651
#17	ARNTL2	27	9	36	1118.776
#18	GATA3	26	14	40	2241.305
#19	ELF4	26	6	32	668.563
#20	GATAD2A	26	7	33	1535.777
#21	NR3C1	26	9	35	2036.408
#22	TFAP2A	26	5	31	822.283
#23	HOXA7	26	9	35	737.42
#24	IRX5	25	12	37	1218.109
#25	CEBPA	25	11	36	2228.891
#26	FOXC1	25	9	34	562.279
#27	RUNX2	25	16	41	3181.587
#28	ETS2	24	7	31	276.478
#29	ELF3	23	13	36	1902.581
#30	HOXA1	23	8	31	453.798
#31	L3MBTL4	23	10	33	1768.864
#32	HOXA5	22	6	28	617.149
#33	ZNF83	22	8	30	1408.809
#34	RORC	22	12	34	2231.711
#35	SNAI2	22	10	32	942.609
#36	HOXA10	22	5	27	164.188
#37	TBX2	22	8	30	1046.873
#38	MEF2C	22	4	26	1123.254
#39	IRF6	22	6	28	1542.477
#40	RUNX3	21	9	30	2124.294
#41	HOXA4	21	6	27	1399.736
#42	GATA6	21	11	32	1827.737
#43	MAFF	21	9	30	595.261
#44	TSC22D3	21	8	29	373.288
#45	FOXF2	21	6	27	99.621
#46	HOXC10	21	7	28	1905.586
#47	NKX3-2	21	11	32	1464.226
#48	TRPS1	20	11	31	965.246
#49	HMGA1	20	8	28	4225.595
#50	HOXB4	20	5	25	948.466
#51	ZNF91	19	13	32	1823.881
#52	ETV5	19	13	32	1759.526
#53	POU2F3	19	8	27	488.431
#54	HHEX	19	10	29	1187.11
#55	TAF4B	19	10	29	1711.46
#56	IRX3	19	11	30	1097.743
#57	CREBZF	19	1	20	63.023
#58	TCF4	19	16	35	2191.088
#59	ENO1	18	7	25	802.022
#60	SCAND2	18	8	26	1870.959
#61	GRHL3	18	14	32	1023.634
#62	ZEB2	18	6	24	438.861
#63	FOXO6	18	7	25	1165.75
#64	EGR4	18	11	29	1551.746
#65	FOXO1	18	13	31	2351.366
#66	ZNF175	18	11	29	2178.013
#67	THRB	18	10	28	2152.24
#68	NKX3-1	18	10	28	1919.978
#69	ETV4	18	16	34	1559.736
#70	EN1	18	11	29	2206.247
#71	EGR3	18	7	25	1280.062
#72	PLAG1	17	12	29	866.089
#73	HOXA11	17	9	26	201.692
#74	PBX4	17	14	31	1616.183
#75	MMP14	17	10	27	1171.586
#76	LZTS1	17	13	30	1307.749
#77	FOXQ1	17	10	27	1159.007
#78	FOXL1	17	6	23	700.497
#79	ESRRG	17	12	29	1514.106
#80	PROX1	17	9	26	1810.865
#81	ZNF256	17	6	23	814.044
#82	SMAD3	16	12	28	693.889
#83	SOX7	16	6	22	329.394
#84	PCGF2	16	10	26	1407.625
#85	IRX2	16	14	30	1253.835
#86	HOXC11	16	11	27	1212.361
#87	PAX3	16	11	27	2597.857
#88	SOX9	16	12	28	2503.614
#89	STAT2	16	9	25	1359.923
#90	C12orf28	16	15	31	1402.683
#91	KLF4	16	8	24	1321.717
#92	TBX3	16	11	27	803.443
#93	KLF6	16	13	29	2181.263
#94	SMAD4	16	7	23	1896.932
#95	PAX8	16	11	27	1260.507
#96	HOXD10	16	6	22	1067.481
#97	PLSCR1	15	13	28	1032.686
#98	NFIA	15	15	30	1796.014
#99	PAX6	15	14	29	596.697
#100	HOXC13	15	12	27	818.781
#101	ZNF165	15	8	23	855.312
#102	ZNF117	15	10	25	1082.946
#103	ZIC2	15	11	26	1761.95
#104	MKX	15	12	27	1544.022
#105	THRA	15	12	27	1522.767
#106	GLIS3	15	14	29	589.944
#107	FOXD1	15	12	27	1162.191
#108	HOXA9	15	14	29	706.903
#109	MEIS2	14	11	25	1037.941
#110	NFE2L2	14	9	23	1261.899
#111	NEUROG1	14	10	24	1489.35
#112	ARX	14	8	22	517.081
#113	MSC	14	9	23	795.556
#114	TRIM22	14	13	27	1126.9
#115	HES5	14	12	26	1223.42
#116	TFEB	14	8	22	1484.764
#117	NKX2-8	14	13	27	1860.084
#118	CREB5	14	11	25	1499.39
#119	HMX2	14	11	25	1653.224
#120	TSC22D2	14	10	24	824.312
#121	EHF	14	16	30	1862.88
#122	ONECUT2	14	11	25	1302.909
#123	DLX2	14	10	24	1134.899
#124	ZNF444	14	9	23	1289.537
#125	SMAD6	14	7	21	791.998
#126	POU6F1	14	12	26	1195.412
#127	NFE2L3	14	10	24	483.587
#128	MAF	14	11	25	1536.08
#129	NPAS2	14	11	25	2000.936
#130	PITX2	13	11	24	1268.687
#131	ZNF24	13	9	22	863.805
#132	SREBF1	13	10	23	635.371
#133	NR4A3	13	11	24	2260.965
#134	GATA4	13	6	19	887.448
#135	TRIM29	13	16	29	1607.499
#136	PBX3	13	8	21	759.533
#137	BHLHE40	13	12	25	1726.439
#138	CITED2	13	12	25	1229.755
#139	MYCN	13	13	26	1156.296
#140	BATF	13	14	27	1525.922
#141	FOXA2	13	5	18	435.817
#142	CBFA2T3	13	12	25	1551.574
#143	NR2F2	13	8	21	1539.604
#144	FOS	13	9	22	830.248
#145	TCF7	13	10	23	966.513
#146	CREB3L1	13	12	25	684.859
#147	EBF1	13	10	23	653.782
#148	TWIST2	13	13	26	1547.549
#149	ANKRD30A	13	14	27	1556.932
#150	VTN	13	13	26	1728.623
#151	DLX3	12	11	23	777.17
#152	ASCL2	12	11	23	626.241
#153	ARNT2	12	11	23	3566.355
#154	CSRNP3	12	9	21	1268.534
#155	IRF9	12	14	26	2272.04
#156	MEIS3P1	12	12	24	1386.981
#157	SCML1	12	7	19	960.757
#158	KCNH8	12	9	21	1108.994
#159	DLX1	12	9	21	1448.288
#160	ZNF238	12	11	23	994.232
#161	BACH2	12	8	20	823.463
#162	SOX13	12	10	22	608.761
#163	RARB	12	11	23	991.584
#164	SOX6	12	10	22	1265.719
#165	NEUROD2	12	8	20	1279.394
#166	FOXO3	12	10	22	1471.861
#167	ETV1	12	15	27	1001.257
#168	TBX5	12	7	19	726.488
#169	RARA	12	13	25	1307.051
#170	TP63	12	14	26	1487.999
#171	SIM1	12	20	32	3431.425
#172	AHR	12	10	22	1051.322
#173	PPARA	12	10	22	690.322
#174	ALX1	12	11	23	1096.333
#175	E2F5	12	13	25	1579.154
#176	LMX1B	12	12	24	1109.674
#177	ZFHX4	12	16	28	1752.451
#178	ISL1	12	9	21	1642.255
#179	EPAS1	12	10	22	1356.393
#180	EMX1	12	12	24	1411.118
#181	NR4A2	12	10	22	1168.984
#182	SMAD9	12	12	24	1195.904
#183	FOXG1	11	11	22	441.067
#184	ZSCAN23	11	11	22	918.242
#185	ESX1	11	10	21	1297.93
#186	HEY1	11	17	28	2531.654
#187	C11orf9	11	10	21	751.313
#188	MEOX1	11	12	23	1067.49
#189	MEIS3	11	14	25	1444.836
#190	ETV6	11	8	19	848.126
#191	NR4A1	11	8	19	405.47
#192	E2F8	11	6	17	797.615
#193	ZHX2	11	12	23	1330.622
#194	BLZF1	11	9	20	859.259
#195	HOXB6	11	4	15	764.608
#196	POU2F1	11	11	22	2976.303
#197	TP73	11	9	20	1238.848
#198	SOX10	11	8	19	710.108
#199	RUNX1	11	12	23	1728.091
#200	NR2F1	11	13	24	1153.067
#201	BTG2	11	12	23	840.76
#202	MSX1	11	19	30	2132.78
#203	HIC1	11	13	24	520.482
#204	POU3F2	11	14	25	1699.514
#205	PGR	11	11	22	578.478
#206	SPIB	11	13	24	943.371
#207	ZNF287	11	11	22	1885.709
#208	GLI3	11	14	25	1996.434
#209	SOX5	11	12	23	1438.092
#210	GATA2	11	10	21	1218.246
#211	JUN	11	12	23	745.407
#212	TFCP2	11	11	22	1463.244
#213	CITED1	11	15	26	1644.409
#214	HOXB9	11	4	15	170.299
#215	MSX2	11	11	22	909.09
#216	MEIS1	11	13	24	1357.463
#217	ZFP42	11	14	25	1800.86
#218	ZNF197	11	9	20	1406.862
#219	HOXB3	11	4	15	575.581
#220	TP53	11	13	24	1879.228
#221	CSDA	11	13	24	1852.65
#222	STAT5B	11	12	23	1284.272
#223	HOXD13	11	12	23	970.895
#224	MAFB	11	19	30	998.13
#225	FOSL2	11	7	18	752.943
#226	KLF16	11	9	20	2364.524
#227	HOXA2	10	15	25	540.346
#228	HR	10	9	19	1030.334
#229	MYC	10	15	25	1180.544
#230	DMRT2	10	11	21	838.434
#231	HOXB2	10	10	20	1001.522
#232	POU3F1	10	11	21	1375.424
#233	SOX8	10	16	26	1546.61
#234	CEBPD	10	16	26	1868.357
#235	ZNF93	10	13	23	1693.69
#236	MESP1	10	10	20	1148.365
#237	RCAN1	10	11	21	1370.834
#238	NR0B1	10	10	20	871.402
#239	ALX3	10	10	20	927.884
#240	ZNF134	10	10	20	916.162
#241	SATB2	10	13	23	980.417
#242	PRRX1	10	5	15	118.654
#243	TFAP2C	10	12	22	422.396
#244	MTA2	10	9	19	885.686
#245	IKZF3	10	13	23	1692.995
#246	TSHZ3	10	14	24	2122.326
#247	LMO4	10	11	21	1093.904
#248	IRF4	10	13	23	979.035
#249	CREBBP	10	8	18	665.521
#250	SP140	10	8	18	552.375
#251	HES6	10	10	20	583.1
#252	HOXD4	10	10	20	1141.194
#253	HOXB8	10	7	17	1296.074
#254	ZFP36L1	10	13	23	1491.41
#255	NPAS1	10	12	22	1127.089
#256	ZNF131	10	9	19	1246.786
#257	KLF5	10	14	24	1836.964
#258	EGR1	10	12	22	819.292
#259	KDM5A	10	14	24	1012.205
#260	ZFHX2	10	8	18	626.381
#261	EN2	10	15	25	1364.905
#262	CDKN2A	10	17	27	3603.309
#263	TEAD2	10	9	19	538.344
#264	MSRB2	9	13	22	1183.236
#265	TWIST1	9	18	27	1078.396
#266	WNT5A	9	13	22	1178.245
#267	PPARG	9	14	23	615.72
#268	HLF	9	15	24	742.703
#269	KLF12	9	14	23	1047.704
#270	ZFPM2	9	13	22	1492.458
#271	SIX3	9	15	24	1268.198
#272	VAV1	9	11	20	1024.637
#273	SOX30	9	13	22	891.73
#274	EGR2	9	9	18	756.851
#275	NR3C2	9	13	22	1142.113
#276	FOXA3	9	13	22	1649.9
#277	ZSCAN12	9	15	24	1651.885
#278	HOXD11	9	11	20	1296.264
#279	TAL1	9	11	20	1082.988
#280	SIX2	9	8	17	932.374
#281	HOXC5	9	14	23	669.488
#282	GSC2	9	12	21	987.352
#283	MLLT10	9	13	22	1207.023
#284	TRERF1	9	13	22	2681.772
#285	PAX4	9	15	24	1492.86
#286	TEAD1	9	9	18	580.576
#287	RERE	9	9	18	1017.5
#288	ZNF3	9	10	19	616.108
#289	POU3F3	9	9	18	508.492
#290	BCL6	9	7	16	613.339
#291	KLF9	9	11	20	878.879
#292	ZHX3	9	14	23	1554.225
#293	KDM5B	9	11	20	891.517
#294	ZXDC	9	13	22	1278.056
#295	MEF2D	9	8	17	757.664
#296	TADA2A	9	13	22	1318.46
#297	ATF6B	9	14	23	1224.222
#298	ZNF18	9	8	17	625.311
#299	HIRA	9	13	22	1857.388
#300	TLX2	9	12	21	1099.17
#301	IRX1	8	12	20	843.18
#302	ZSCAN20	8	13	21	1279.878
#303	PKNOX2	8	10	18	602.933
#304	NKX6-2	8	15	23	1060.38
#305	FOXK2	8	9	17	528.666
#306	GATA5	8	11	19	780.98
#307	NEUROD1	8	14	22	1085.136
#308	OLIG2	8	13	21	1087.656
#309	NFATC1	8	19	27	1797.237
#310	ZNF135	8	7	15	820.69
#311	RFX3	8	9	17	582.163
#312	FOXC2	8	8	16	444.864
#313	MLL	8	14	22	1157.29
#314	NFYA	8	11	19	873.632
#315	HDX	8	14	22	1437.711
#316	MBD1	8	10	18	633.358
#317	PGBD1	8	14	22	815.208
#318	GBX2	8	12	20	1077.343
#319	RCOR1	8	12	20	1104.265
#320	HMG20B	8	10	18	542.291
#321	HIF3A	8	11	19	696.938
#322	GATAD2B	8	9	17	655.605
#323	SALL1	8	11	19	928.128
#324	AFF4	8	8	16	469.358
#325	MNX1	8	8	16	552.462
#326	NFIX	8	16	24	1285.184
#327	RUNX1T1	8	16	24	1824.93
#328	TFDP2	8	11	19	955.966
#329	MYT1	8	11	19	1182.866
#330	NFE2	8	10	18	776.019
#331	HOXB13	8	15	23	2094.017
#332	OVOL1	8	9	17	515.751
#333	SLC26A3	8	12	20	1296.431
#334	NFKB2	8	10	18	982.221
#335	POU4F1	8	10	18	462.043
#336	SMAD2	8	9	17	893.903
#337	ZNF80	8	12	20	1462.216
#338	NANOGNB	8	9	17	710.646
#339	NFAT5	7	10	17	920.317
#340	TCF12	7	16	23	1244.191
#341	FOXP4	7	11	18	795.837
#342	HMX1	7	13	20	900.888
#343	ELF5	7	20	27	1467.945
#344	LHX6	7	11	18	685.934
#345	ZKSCAN3	7	11	18	554.848
#346	ZIC1	7	16	23	1134.273
#347	BHLHE41	7	15	22	582.202
#348	IRX4	7	16	23	937.734
#349	IRF7	7	11	18	936.407
#350	ERG	7	14	21	1291.266
#351	SOX4	7	11	18	784.494
#352	SOX17	7	10	17	931.188
#353	LHX9	7	12	19	674.213
#354	TCF7L1	7	10	17	797.985
#355	SOX11	7	15	22	1237.88
#356	NFIB	7	16	23	1204.063
#357	BCL11B	7	12	19	759.614
#358	DMBX1	7	9	16	699.861
#359	ZNF639	7	3	10	195.698
#360	TFCP2L1	7	19	26	1551.193
#361	ISL2	7	12	19	1290.137
#362	SALL2	7	12	19	898.06
#363	RARG	7	12	19	755.713
#364	MLXIPL	7	8	15	620.955
#365	FOXJ1	7	13	20	1651.369
#366	TBX20	7	15	22	1275.594
#367	ZSCAN18	7	13	20	1010.378
#368	MYNN	7	12	19	999.911
#369	TLX3	7	11	18	788.524
#370	ZNF496	7	10	17	633.553
#371	ZNF167	7	15	22	868.145
#372	CUX2	7	11	18	669.614
#373	HNF4G	7	21	28	1327.229
#374	ATF2	7	8	15	1727.156
#375	OTX2	7	13	20	1117.192
#376	IRF1	7	15	22	1070.618
#377	TADA3	7	11	18	717.713
#378	FOXP1	7	11	18	665.818
#379	FOXL2	7	13	20	1084.326
#380	GLI2	7	9	16	277.676
#381	RORA	7	13	20	1765.891
#382	NR5A2	7	17	24	924.089
#383	NR1H4	7	13	20	3254.123
#384	ZNF397	7	12	19	773.375
#385	ZIC5	7	12	19	1279.352
#386	LHX1	7	14	21	657.404
#387	ST18	6	12	18	733.649
#388	MGA	6	11	17	985.266
#389	SIX1	6	13	19	871.288
#390	ATOH1	6	8	14	844.088
#391	HOXC12	6	10	16	500.446
#392	FLI1	6	12	18	797.227
#393	PTH	6	11	17	736.494
#394	NKX2-1	6	13	19	624.975
#395	PRDM2	6	15	21	1053.366
#396	SOX1	6	15	21	1312.812
#397	ZSCAN2	6	10	16	407.666
#398	FOXE1	6	16	22	615.299
#399	FUBP1	6	8	14	354.896
#400	STAT6	6	17	23	1179.206
#401	ZIC3	6	9	15	886.816
#402	MEOX2	6	18	24	1256.579
#403	ZFHX3	6	13	19	1130.681
#404	ZNF81	6	13	19	655.21
#405	VSX1	6	11	17	505.789
#406	SOX18	6	13	19	1308.648
#407	HOXD8	6	6	12	328.023
#408	LHX8	6	15	21	1399.399
#409	ESRRB	6	12	18	548.858
#410	NKX2-5	6	16	22	494.026
#411	TFEC	6	12	18	683.544
#412	NFATC2	6	16	22	634.181
#413	ZNF483	6	12	18	823.172
#414	MITF	6	14	20	1105.67
#415	TCF7L2	6	14	20	853.436
#416	HOXA13	6	18	24	1182.988
#417	SIN3A	6	12	18	866.288
#418	MECOM	6	12	18	555.998
#419	RB1	6	6	12	672.031
#420	STRN3	6	14	20	1195.876
#421	NPAT	6	9	15	480.257
#422	KLF15	6	13	19	964.926
#423	ZNF323	6	10	16	629.56
#424	SOX21	5	14	19	801.821
#425	KLF2	5	15	20	750.583
#426	PPARD	5	10	15	562.137
#427	NKX2-3	5	11	16	534.021
#428	ETV7	5	13	18	1204.041
#429	ZNF90	5	14	19	611.259
#430	FOSB	5	10	15	737.363
#431	MTA3	5	11	16	937.282
#432	ELK4	5	10	15	595.904
#433	ZNF215	5	11	16	575.941
#434	POU1F1	5	12	17	924.735
#435	FOXI1	5	11	16	1006.19
#436	ZSCAN4	5	9	14	451.742
#437	CREM	5	14	19	658.784
#438	HNRNPAB	5	15	20	911.288
#439	NRL	5	12	17	1144.547
#440	HOXD3	5	9	14	296.636
#441	HOPX	5	14	19	563.722
#442	HESX1	5	13	18	730.723
#443	CREB3L3	5	13	18	790.944
#444	HOXB7	5	8	13	365.12
#445	GAS7	5	14	19	679.767
#446	BNC1	5	7	12	252.389
#447	SMAD1	5	10	15	766.213
#448	PEG3	5	13	18	633.217
#449	PRRX2	5	14	19	769.185
#450	SCAND1	5	11	16	530.274
#451	ZNF396	5	13	18	507.188
#452	SIM2	5	16	21	511.527
#453	HSF5	5	10	15	1206.167
#454	FOXP3	5	10	15	824.644
#455	ALS2CR8	5	12	17	661.888
#456	NANOG	5	14	19	842.93
#457	TULP4	5	18	23	1109.961
#458	GSX1	4	12	16	503.003
#459	MESP2	4	11	15	767.901
#460	TBX19	4	12	16	385.898
#461	ONECUT1	4	13	17	674.599
#462	TSHZ2	4	10	14	641.292
#463	LMX1A	4	11	15	451.575
#464	SOX15	4	9	13	307.717
#465	BARX1	4	14	18	769.99
#466	LASS5	4	12	16	781.48
#467	TBX22	4	13	17	961.23
#468	MAFG	4	13	17	509.173
#469	SPIC	4	13	17	303.667
#470	FOXP2	4	14	18	675.399
#471	ZNF132	4	14	18	1398.019
#472	HEY2	4	18	22	541.119
#473	SOX2	4	18	22	1002.632
#474	ZNF157	4	15	19	784.786
#475	NFATC4	4	13	17	1349.66
#476	ZNF85	4	14	18	830.537
#477	TFAP2B	4	16	20	512.677
#478	ZNF207	4	11	15	291.713
#479	REST	4	8	12	296.366
#480	TBX1	4	15	19	689.998
#481	SATB1	3	15	18	562.627
#482	MXD1	3	15	18	789.506
#483	TAF4	3	12	15	590.42
#484	ASCL1	3	15	18	372.76
#485	TEF	3	9	12	350.027
#486	SNAI3	3	10	13	538.795
#487	RFXAP	3	14	17	834.489
#488	LEF1	3	24	27	615.921
#489	IKZF1	3	11	14	401.804
#490	MYT1L	3	10	13	235.288
#491	FOXD3	3	16	19	608.783
#492	MYF5	3	10	13	321.271
#493	C2orf3	3	9	12	584.29
#494	RORB	3	10	13	713.203
#495	CTBP1	2	13	15	384.819
#496	PITX1	2	14	16	202.641
#497	TCF21	2	11	13	248.528
#498	GCM1	2	9	11	196.335
#499	LHX2	1	16	17	117.994
#500	WT1	1	12	13	230.047